As part of the audition for the role of the Czech émigré who restores the artistic fire in a heartbroken Irish busker in the musical "Once," Milioti was given a couple of weeks to learn to play a classical piano piece. She threw herself into the task with discipline and rigor and won the role of "the Girl." For her efforts, she has earned enthusiastic reviews and a Tony nomination for lead actress in a musical. Milioti has yet to see the film with Markéta Irglová and Glen Hansard on which the musical is based, but she captures its pathos and romanticism.
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